Curriculum & Courses
M.S. in Ecology
Required Courses: 6 credits
Fundamentals Courses: 6 credits
Ecology Tools Courses: 3 credits
Additional Electives, Independent Study, Research, and Dissertation: Minimum 15 credits
Total: Minimum 30 credits
Ph.D. in Ecology
Required Courses: 8 credits
Fundamentals Courses: 6 credits
Ecology Tools Courses: 3 credits
Additional Electives, Independent Study, Research, and Dissertation: Minimum 55 credits
Total: Minimum 72 credits
Ph.D. in Ecology -
HEI Specialization
Required Courses: 11 credits
Fundamentals Courses: 6 credits
HEI Fundamentals Courses: 3 credits
Ecology Tools Courses: 3 credits
Qualitative HEI Tools: 3 credits
Additional Electives, Independent Study, Research, and Dissertation: Minimum 46 credits
Total: Minimum 72 credits
Coursework Information
The GDPE curriculum is designed to balance providing the fundamentals of ecology that unite students in our program with allowing customization for each student’s individualized scholarly pursuits.
Required Courses (MS – 6 credits, PhD – 8 credits)
These required courses provide the common foundation for all GDPE students and ensure that all students engage in key intellectual opportunities within the program. The many different electives available allow students to specialize and customize their individual MS or PhD degree.
| Course | Title | Credits | Semester |
|---|---|---|---|
| ECOL 505 | Foundations of Ecology | 3 | Fall |
| ECOL 571 | Distinguished Ecologists Lecture Series (MS – 1 credit, PhD – 2 credits) | 1-2 | Spring |
| ECOL 592 | Interdisciplinary Seminars in Ecology (MS – 1 course, PhD – 2 courses) | 1-3 | Fall & Spring |
| ECOL 693 | Research Seminars for Professional Communication (Section 1 – oral, Section 2 – written) | 1 | Spring |
| ANTH 530 (required for HEI specialization only) | Human-Environment Interactions | 3 | Fall |
Ecology Fundamentals (MS & PhD – 6 credits)
| Course | Title | Credits | Semester |
|---|---|---|---|
| BZ 525 | Advanced Conservation & Evolutionary Genomics | 4 | Fall (odd years) |
| BZ 526/BSPM 526 | Evolutionary Ecology | 3 | Fall (odd years) |
| BZ 535 | Behavioral Ecology | 3 | Spring (odd years) |
| BZ 548 | Theory of Population and Evolutionary Ecology | 4 | Fall |
| ECOL 600 | Community Ecology | 3 | Spring |
| ECOL 610 | Ecosystem Ecology | 3 | Fall |
| ECOL 620 | Applications in Landscape Ecology | 4 | Spring (odd years) |
| ESS 660 | Biogeochemical Cycling | 3 | Spring |
| ESS 575 | Models for Ecological Data | 4 | Fall |
| FW 662 | Wildlife Population Dynamics | 3 | Spring (odd years) |
| HORT 576 | Advanced Environmental Plant Stress Physiology | 4 | Spring (odd years) |
Ecology Tools (MS & PhD – 3 credits)
| Course | Title | Credits | Semester |
|---|---|---|---|
| AREC 535 | Applied Econometrics | 3 | Fall |
| AREC 635 | Econometric Theory I | 3 | Fall |
| AREC 735 | Econometric Theory II | 3 | Spring |
| BZ 548 | Theory of Population and Evolutionary Ecology | 4 | Spring |
| BZ 577/MIP 577 | Computer Analysis in Population Genetics | 2 | Fall |
| CIVE/WR 524 | Modeling Watershed Hydrology | 3 | Spring |
| ERHS 544/STAT 544 | Biostatistical Methods for Quantitative Data | 3 | Spring |
| ESS 655 | Multivariate Analysis for Community Ecology | 2 | Fall (even years) |
| ESS 565 | Niche Models | 4 | Fall (odd years) |
| ESS 575 | Models for Ecological Data | 4 | Fall |
| FW 551 | Design of Fish and Wildlife Studies | 3 | Fall |
| FW 552 | Applied Sampling for Wildlife/Fish Studies | 3 | Spring (odd years) |
| FW 663 | Sampling and Analysis of Vertebrate Populations | 5 | Spring (even years) |
| FW 673/STAT 673 | Hierarchical Modeling in Ecology | 3 | Fall (odd years) |
| GR 503/NR 503 | Remote Sensing and Image Analysis | 4 | Fall |
| MATH 530 | Mathematics for Scientists and Engineers | 4 | Fall |
| MATH 540 | Dynamical Systems | 3 | Fall |
| NR 505 | Concepts in GIS | 4 | Fall |
| NR 506 | GIS Methods for Resource Management | 4 | Spring |
| NR 512 | Spatial Statistical Modeling-Natural Resources | 3 | Fall |
| NR 523/STAT 523 | Quantitative Spatial Analysis | 3 | Spring |
| NR 554/ANTH 554 | Ecological and Social Agent-based Modeling | 3 | Spring (odd years) |
| NRRT 765 | Applied Multivariate Analysis | 3 | Fall |
| SOCR 522 | Micrometeorology | 3 | Spring |
| SOCR 620 | Modeling Ecosystem Biogeochemistry | 3 | Fall (odd years) |
| SOCR 670 | Terrestrial Ecosystems Isotope Ecology | 3 | Spring (even years) |
| STAR 511 | Design and Data Analysis for Researchers I | 4 | Fall, Spring, Summer |
| STAR 512 | Design and Data Analysis for Researchers II | 4 | Spring |
| STAT 520 | Introduction to Probability Theory | 4 | Fall |
| STAT 521 | Stochastic Processes I | 3 | Spring |
| STAT 530 | Mathematical Statistics | 3 | Spring |
| STAT 540 | Data Analysis and Regression | 3 | Fall |
| STAT 560 | Applied Multivariate Analysis | 3 | Fall, Spring |
| STAT 675A | Topics in Statistical Methods: Sampling | 1-3 | Fall, Spring, Summer |
| WR 674 | Data Issues in Hydrology | 3 | Spring (even years) |
Additional Electives (MS – 15 credits, PhD – 55 credits)
This list covers many, but not all, courses likely to be relevant to our diverse discipline. Note that any “Fundamental” or “Tools” course can be used as an elective if it has not been used to fulfill another requirement| Course | Title | Credits | Semester |
|---|---|---|---|
| ANTH 530 | Human Environment Interactions | 3 | Fall (odd years) |
| AREC 505 | Agricultural Production Economics | 3 | Fall |
| AREC 535 | Applied Econometrics | 3 | Fall |
| AREC 540 | Economics of Natural Resources | 3 | Spring |
| AREC 541 | Environmental Economics | 3 | Spring |
| AREC 635 | Econometric Theory I | 3 | Fall |
| ATS 760 | Global Carbon Cycle | 2 | Spring (odd years) |
| BSPM 523 | Evolution and Classification of Insects | 4 | Fall (odd years) |
| BSPM 528 | Invasive Plants and Weeds: Ecosystems to Molecules | 3 | Spring (even years) |
| BZ 561 | Landscape Ecology | 3 | Fall |
| BZ 505 | Cognitive Ecology | 3 | Fall (odd years) |
| BZ 510 | Zoophysiological Ecology | 3 | Spring (odd years) |
| BZ 525 | Molecular Ecology | 4 | Fall (odd years) |
| BZ 526/BSPM 526 | Evolutionary Ecology | 3 | Fall (odd years) |
| BZ 530 | Ecological Plant Morphology | 2 | Spring (even years) |
| BZ 535 | Behavioral Ecology | 3 | Fall (even years) |
| BZ 548 | Theory of Population and Evolutionary Ecology | 4 | Fall |
| BZ 555 | Reproductive Biology of Higher Plants | 3 | Spring (odd years) |
| BZ 572 | Phytoremediation | 3 | Fall (odd years) |
| BZ 568/FW 568 | Sustaining River Biodiversity and Ecosystem Services in a Dynamically Changing World | 3 | Spring |
| ECOL 600 | Population and Community Ecology | 3 | Spring |
| ECOL 610 | Ecosystem Ecology | 3 | Fall (even years) |
| ECOL 620 | Applications in Landscape Ecology | 4 | Spring (odd years) |
| ESS 660 | Biogeochemical Cycling | 3 | Spring |
| ESS 524 | Foundations for Greenhouse Gas Accounting and Management | 3 | Fall |
| F 610 | Advance Forest Ecology | 3 | Spring (odd years) |
| F 624 | Fire Ecology | 3 | Fall, Spring |
| FW 544 | Ecotoxicology | 3 | Spring |
| FW 555 | Conservation Biology | 3 | Spring |
| FW 558 | Conservation Genetics of Wild Populations (on-line) | 3 | Spring |
| FW 567 | Wildlife Disease Ecology | 3 | Fall (odd years) |
| FW 605 | Advanced Physiological Ecology of Fishes | 4 | Spring (even years) |
| FW 662 | Wildlife Population Dynamics | 3 | Spring (odd years) |
| GEOL 652 | Fluvial Geomorphology | 3 | Fall |
| GEOL 551 | Groundwater Modeling | 3 | Spring |
| GES 520 | Issues in Global Environmental Sustainability | 3 | Fall, Spring |
| HORT 571 | Soil Plant Water Relations | 3 | Spring (even years) |
| HORT 576 | Advanced Environmental Plant Stress Physiology | 4 | Spring (odd years) |
| MIP 624 | Advanced Topics in Microbial Ecology | 2 | Fall (odd years) |
| MIP 570 | Functional Genomics | 3 | Fall |
| NR 504 | Computer Analysis of Remote Sensing Data | 4 | Spring |
| NR 510 | Ecosystem Services: Theory and Practice | 3 | Spring |
| NR 625 | Community-Based Natural Resource Management | 3 | Spring |
| NR 578 | Ecology of Disturbed Lands | 3 | Spring |
| NR 678 | Advanced Ecological Restoration | 4 | Fall, Spring |
| RS 565 | Riparian Ecology and Management | 3 | Spring |
| RS 630 | Ecology of Grasslands and Shrublands | 3 | Spring |
| SOCR 540 | Soil-Plant-Nutrient Relationships | 3 | Spring (even years) |
| SOCR 550 | Advanced Soil Genesis | 3 | Spring (even years) |
| SOCR 567 | Environmental Soil Chemistry | 4 | Spring |
| SOCR 755 | Advanced Soil Microbiology | 3 | Spring (even years) |
| SOCR 770 | Advanced Soil Physics | 4 | Spring (even years) |
| STAR 500-549* | Applied Stats for Researchers (STAR) | Varies | |
| WR 574 | Advanced Snow Hydrology | 4 | Fall (even years) |
| WR 575 | Snow Hydrology Field Methods | 1 | Spring |
| ECOL 581A2/AB 581A2 | Insect Ecology | 3 | Spring (odd years) |
Human-Environment Interactions Fundamentals (PhD-HEI – 3 credits)
| Course | Title | Credits | Semester |
|---|---|---|---|
| ANTH 515 | Culture and Environment | 3 | Fall |
| ANTH 529 | Anthropology and Sustainable Development | 3 | Fall |
| ANTH 535 | Globalization and Culture Change | 3 | Fall (even years) |
| ANTH 540 | Medical Anthropology | 3 | Spring (even years) |
| ANTH 545 | Global Mental Health–Theory and Method | 3 | Fall (odd years) |
| ANTH 555 | Paleoindian Archaeology | 3 | Fall (even years) |
| ANTH 571 | Anthropology and Global Health | 3 | Fall (even years) |
| ANTH 572 | Human Origins | 3 | Spring (odd years) |
| NR 625 | Community-Based Natural Resource Management | 3 | Spring |
| POLS 739 | International Environmental Politics | 3 | Fall, Spring |
| SOC 668 | Environmental Sociology | 3 | Fall |
Qualitative Ecology Tools (PhD-HEI 3 credits)
| Course | Title | Credits | Semester |
|---|---|---|---|
| EDRM 704 | Qualitative Research | 3 | Fall, Summer |
| EDRM 706 | Analysis of Variance–Education Research | 3 | Spring, Summer |
| NRRT 665 | Survey Research and Analysis | 3 | Spring |
| POLS 621 | Qualitative Methods in Political Science | 3 | Spring (odd years) |
| SOC 610 | Seminar in Methods of Qualitative Analysis | 3 | Spring (even years) |
Adding non-regular credits can be confusing! Watch this helpful walk-through on How to add Independent Study/Research/Thesis/Dissertation Credits
| Course | Course Title | Term | Course Description | |
|---|---|---|---|---|
| ECOL 505 | Foundations of Ecology | Fall | Overview of the science of ecology; what questions are asked, how they are answered. Prerequisites: One course in ecology CRN: 60162 Credits: 3 | |
| ECOL 571 | Advanced Topics in Ecology Distinguished Ecologists Lecture Series | Spring | Current research topics presented and analyzed by visiting scientists. Attend each Distinguished Ecologist seminar during the semester for which you are registered. This course does not meet every week. Prerequisites: One course in ecological principles. Section 001 – MS students. CRN: 18219 Credits: 1; Section 002 – PhD students. CRN: 18220 Credits: 2; Section 003 (intermittently offered; verify prior to registration) – PhD students. – CRN: 21250 Credits: 2 | |
| ECOL 592 | Interdisciplinary Seminars in Ecology | Fall/Spring | Concepts and principles of basic and applied ecology in an interdisciplinary context. Specific topics vary. Prerequisites: One 300- or 400-level course in ecology. Course details and CRNs: ECOL592 Interdisciplinary Seminar Credits: 1 – 3 (vary based on section) | |
| ECOL 600 | Community Ecology Lecture | Spring | Current theories on the dynamics and regulation of populations and communities of organisms. Must enroll in a recitation section. Prerequisites: One course each on general ecology, calculus, and statistics. CRN: 20590 Credits: 3 Recitation: Section R01 – CRN: 20591; Section R02 – CRN: 20592 | |
| ECOL 610 | Ecosystem Ecology Lecture | Fall | Concepts, methods, issues in ecosystem science: energy and matter cycling, systems perspectives, simulation modeling, sustainability, global change. Must enroll in a recitation section. Prerequisites: LIFE 320 or any ECOL course. CRN: 66480 Credits: 3 Recitation: Section R01 – CRN: 66481 | |
| ECOL 620 | Applications in Landscape Ecology Lecture | Spring (Odd years only) | Spatial patterning of landscape elements and dynamics of ecological systems; spatial heterogeneity. Influence on biotic and abiotic processes. Prerequisites: Previous coursework in geographic information systems, ecology, statistics, and mathematics. CRN: 66478 Credits: 2 | |
| ECOL 693 | Research Seminars for Professional Communication (written or oral) | Spring | Critique of research programs, plans, and ecological theory. Prerequisites: Written consent of instructor. Section 001 – Enhancing oral presentation skills. CRN: 18231; Section 002 – Enhancing proposal writing skills. CRN: 18232; Section 003 (intermittently offered; verify prior to registration) – Enhancing proposal writing skills. CRN: 19376 Credits: 1 | |
| ECOL 695 | Independent Study | Fall/Spring/Summer | Requires a contract, thoroughly outlining the scope of the project, signed by student, instructor and and approved by the Director. Application accessible on the GDPE Forms webpage. Prerequisites: GDPE Approval Fall Semester CRN: 60575 Spring Semester CRN: 10371 Summer Semester CRN: 50821 Credits: 1- 18 | |
| ECOL 698 | Research | Fall/Spring/Summer | Fall Semester CRN: 60576 Spring Semester CRN: 10372 Summer Semester CRN: 50822 Credits: 1- 18 | |
| ECOL 699 | Thesis | Fall/Spring/Summer | Fall Semester CRN: 60277 Spring Semester CRN: 10373 Summer Semester CRN: 50823 Credits: 1- 18 | |
| ECOL 799 | Dissertation | Fall/Spring/Summer | Fall Semester CRN: 60578 Spring Semester CRN: 10374 Summer Semester CRN: 50824 Credits: 1- 18 | |
| Continuous Registration | Fall/Spring/Summer | Fall Semester CRN: 62570 Spring Semester CRN: 12719 Summer Semester CRN: 45836 Credits: n/a Further details on when to register for CR can be found at the Graduate School website. |
ECOL 592 Interdisciplinary Seminar
ECOL592 seminars are designed to cover concepts and principles of basic and applied ecology in an interdisciplinary context. One of the unique things about these seminars is that the topics are driven by faculty & student interests. This means topics change each semester, so be sure to check the ECOL 592 page for current listings. You can also review past seminar topics here.
Requirements:
- Master students are required to take 1 ECOL592 seminar.
- PhD students are required to take 2 separate ECOL592 seminars.
Expectations for Seminars:
Most seminars are 1 credit and meet once a week. Students should expect to have 1-2 hours of preparation time outside of the normal meeting time.
Past seminars have included journal-club-styles, topic-specific discussions with international experts via video conferencing, data analysis and manuscript preparation, and more.
The GDPE conference room can be reserved for ECOL592 meeting times. Email ecology@colostate.edu to organize the reservation
Proposing a seminar topic:
Proposals are not lengthy, but should include details on the format of the seminar, expectations, grading, and outcomes & objectives. If you have questions, please contact the GDPE Assistant Director. Please schedule the seminar time so it does not conflict with DE seminars (Wednesdays at 4pm).
Faculty: submit a proposal through the GDPE website portal here.
Students:
- If you are interested in having a faculty member lead an ECOL592 on a topic that interests you, please reach out to them and encourage them to submit a proposal.
- If you have already completed at least one ECOL592 seminar as a student, you may submit a proposal to co-teach a seminar with a faculty member. Work together, and then submit the proposal through the GDPE website portal here.
Note: For PhD students, you may petition to have your second ECOL592 requirement waived at the end of the semester in which you teach. To do so, please complete the petition through the GDPE web portal here. Please include a letter from your faculty co-instructor confirming your participation in the course. If this is your advisor, they can include that in their supporting letter.
Curriculum information is also available in the CSU Catalog.
To see when the courses are being offered, use the Class Schedule.
If you have questions about specific non-ECOL courses listed above, please reach out to the department or instructor directly.
Credit Requirement Information
Full-Time Graduate Student Credits
Graduate students must be registered for 9 credits or more during fall and spring semesters, or 5 credits or more during summer semester, to be considered a full-time student. Students enrolled in half-time (5 or 6 credits) or more participate in CSU’s Health Care program.
GTA & GRA Credit Requirement
Graduate Teaching Assistantships (GTAs) and Graduate Research Assistantships (GRAs) can only be awarded to students enrolled in at least one on-campus credit during the spring and fall semesters. Continuous Registration (CR, see below) is NOT acceptable to meet the minimum enrollment requirement for a GTA or GRA.
Continuous Registration (CR)
All students admitted to a graduate degree program at CSU are required to be continuously enrolled in their degree programs in fall and spring semesters. Thus, if coursework and research are essentially completed, and a student is working on their thesis or dissertation, they are required to register for at least one credit. This can be done by registering for one credit of coursework or research or through Continuous Registration (CR).
(Fall): CRN #67542
(Spring): CRN #12719
(Summer): CRN #45836
Students registering with CR will be assessed a fee for each semester of CR registration.
Students enrolled with CR in any term are not considered enrolled full time for the purposes of financial aid, Visas, employment, etc. To receive full privileges for the summer semester, students must be enrolled either in the summer or the following fall semester. This policy applies from the first enrollment semester through the graduation term.
Students should contact their advisor if they do not plan to register.
Independent Study, Research, Thesis, and Dissertation Credits
Independent Study (ECOL 695), Research (ECOL 698), Thesis (ECOL 699), and Dissertation (ECOL 799) registration is for non-regular defined, variable-credit courses. Credit hours are determined using a base rate of 48 hours of student effort per credit hour over a 16-week semester. The total number of hours are divided by 48 and the result quotient (rounded to a whole number) will be the number of credits to be used. Speak with your advisor to determine how many research credits should be registered for. Your advisor will assign a grade at the end of the term, which is often done on a S/U (satisfactory/unsatisfactory) basis.
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